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{{cite journal
| author=Gao, F., Bailes, E., Robertson, D. L., Chen, Y., Rodenburg, C. M., Michael, S. F., Cummins, L. B., Arthur, L. O., Peeters, M., Shaw, G. M., Sharp, P. M. and Hahn, B. H. | title=Origin of HIV-1 in the Chimpanzee Pan troglodytes troglodytes | journal=Nature | year=1999 | pages=436-441 | volume=397 | issue=6718 | id={{PMID |9989410}}